Kyrgyzstan July 2008 Zabelia corymbosa quest

Voucher: KYRG 149 

Xanthium spinosum L.

Region: Jalal-Abad Oblast  Ala-Buka Rayon
Gazetteer: west of Joy-Belent 
Coordinates: 41º23'48'' N, 71º24'11'' E  Alt: 1221m 

Spreading small annual plant. Leaves three lobed with a tripartite spine in the axil. Female capitula covered with hooked spines. Male capitula terminal. Ruderal plant.
Dry hill along the road with Alhagi, Echinops, Chenopodiaceae, Caragana.

Identification Notes: Verified with Flora of the USSR. Vol.25, V. L. Komarov. Leaves ovate lobed, stiff hairy adaxially, covered in white hairs abaxially. Spines tripartite yellow at the base of each leaf. Male capitula grouped at apex of stems, florets with valvate corolla, anthers fused at base but then free not forming a column. Female capitula with 2 florets inside, phyllaries fused woody and covered with strong hooked yellow spines with a tiny hole at the apex for the styles to come through. Stigmas linear. Cypsela with straight aculeiform small beak at apex. 
Ancillary Collections: photos 

Landrein, S. 149                8 July 2008

morphological details

male capitulum male capitulum staminal column staminal column male floret male floret

female capitulum female capitulum

phyllaries hooks phyllaries hooks

section of the female capitulum section of the female capitulum

cypsela cypsela

cyspela apex cyspela apex

section of the cypsela section of the cypsela

spines spines

adaxial leaf surface adaxial leaf surface

abaxial leaf surface abaxial leaf surface

field picture
flowering plant
Herbarium voucher
Herbarium voucher

Other field pictures

capitula capitula
capitula capitula